Table 4. Physical environment parameters at the sampling sites (mean ± S.D)

Site	Substrate(%)*					Water velocity (cm/sec)	Water depth (cm)	Water temperature (°C)	Altitude (m)
	B	C	P	G	S
St. 1	11	37	21	10	21	34.4 ± 13.8	13.9 ± 7.7	16.5 ± 5.2	169
St. 2	11	26	15	17	31	32.3 ± 3.2	20.5 ± 7.4	17.7 ± 5.7	98
St. 3	23	24	19	10	24	54.9 ± 24.1	21.8 ± 2.5	17.8 ± 5.9	113
St. 4	0	25	29	17	29	27.6 ± 25.0	11.7 ± 6.1	17.1 ± 6.6	113
St. 5	14	28	20	15	23	47.9 ± 34.4	10.2 ± 2.2	16.9 ± 6.2	68

B: boulder (>256mm), C: cobble (64∼256mm), P: pebble (16∼64mm), G: gravel (2∼16mm), S: sand and silt (≤2mm).